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<article article-type="research-article" dtd-version="1.3" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xml:lang="ru"><front><journal-meta><journal-id journal-id-type="publisher-id">ecodag</journal-id><journal-title-group><journal-title xml:lang="ru">Юг России: экология, развитие</journal-title><trans-title-group xml:lang="en"><trans-title>South of Russia: ecology, development</trans-title></trans-title-group></journal-title-group><issn pub-type="ppub">1992-1098</issn><issn pub-type="epub">2413-0958</issn><publisher><publisher-name>State Institute of Applied Ecology of the Republic of Dagestan</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="doi">10.18470/1992-1098-2016-3-35-89</article-id><article-id custom-type="elpub" pub-id-type="custom">ecodag-856</article-id><article-categories><subj-group subj-group-type="heading"><subject>Research Article</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="ru"><subject>ЭКОЛОГИЯ ЖИВОТНЫХ</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="en"><subject>ECOLOGY OF ANIMALS</subject></subj-group></article-categories><title-group><article-title>ГЕОГРАФИЧЕСКИЕ СВЯЗИ ЖУКОВ-ЧЕРНОТЕЛОК (COLEOPTERA: TENEBRIONIDAE) ТЕТИЙСКОЙ ПУСТЫННО-СТЕПНОЙ ОБЛАСТИ ПАЛЕАРКТИКИ С ИСТОРИЧЕСКИМ ОБЗОРОМ</article-title><trans-title-group xml:lang="en"><trans-title>GEOGRAPHICAL RELATIONS OF DARKLING BEETLES (COLEOPTERA: TENEBRIONIDAE) OF THE PALAEARCTIC TETHYS DESERT-STEPPE REGION WITH THE HISTORICAL REVIEW</trans-title></trans-title-group></title-group><contrib-group><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Абдурахманов</surname><given-names>Г. М.</given-names></name><name name-style="western" xml:lang="en"><surname>Abdurakhmanov</surname><given-names>G. M.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Гайирбег М. Абдурахманов - академик РЭА, доктор биологических наук, профессор, заслуженный деятель науки РФ, заведующий кафедрой биологии и биоразнообразия.</p><p>Ул. Дахадаева, 21, Махачкала, 367001 Россия, e-mail: abgairbeg@rambler.ru</p></bio><bio xml:lang="en"><p>Gayirbeg M. Abdurakhmanov - Academician of Russian Academy of Ecology, Doctor of Biological Sciences, Professor, Honored Scientist of Russia, Head of the department of biology and biodiversity.</p><p>21   Dakhadaeva   st.,   Makhachkala,   367001 Russia. e-mail: abgairbeg@rambler.ru </p></bio><email xlink:type="simple">abgairbeg@rambler.ru</email><xref ref-type="aff" rid="aff-1"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Набоженко</surname><given-names>М. В.</given-names></name><name name-style="western" xml:lang="en"><surname>Nabozhenko</surname><given-names>M. V.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Максим В. Набоженко - кандидат биологических наук, ведущий научный сотрудник ПИБР ДНЦ РАН и доцент кафедры биологии и биоразнообразия </p></bio><bio xml:lang="en"><p>Maxim V. Nabozhenko - PhD, leading scientific researcher  of  Caspian  Institute  of  Biological  Resouces RAS, associate professor of the department of biology and biodiversity</p></bio><xref ref-type="aff" rid="aff-1"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Абдурахманов</surname><given-names>А. Г.</given-names></name><name name-style="western" xml:lang="en"><surname>Abdurakhmanov</surname><given-names>A. G.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Абдурахман  Г.  Абдурахманов  - членкорреспондент РЭА, кандидат биологических наук, доцент кафедры рекреационной географии и устойчивого развития</p></bio><bio xml:lang="en"><p>Abdurakhman G. Abdurakhmanov - corresponding member of REA, candidate of biological sciences, associate professor of the department of recreation geography and sustainable development</p></bio><xref ref-type="aff" rid="aff-2"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Иванушенко</surname><given-names>Ю. Ю.</given-names></name><name name-style="western" xml:lang="en"><surname>Ivanushenko</surname><given-names>Yu. Yu.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Юлия Ю. Иванушенко - аспирант кафедры биологии и биоразнообразия</p></bio><bio xml:lang="en"><p>Yuliya Yu Ivanushenko - postgraduate of Department of biology and biodiversity</p></bio><xref ref-type="aff" rid="aff-2"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Даудова</surname><given-names>М. Г.</given-names></name><name name-style="western" xml:lang="en"><surname>Daudova</surname><given-names>M. G.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Мадина   Г.   Даудова - член-корреспондент   РЭА, кандидат биологических наук, старший преподаватель кафедры биологии и биоразнообразия</p></bio><bio xml:lang="en"><p>Madina G. Daudova corresponding member of REA, candidate of biological sciences, department of biology and biodiversity</p></bio><xref ref-type="aff" rid="aff-2"/></contrib></contrib-group><aff-alternatives id="aff-1"><aff xml:lang="ru">Институт экологии и устойчивого развития, Дагестанский государственный университет, Махачкала; Прикаспийский институт биологических ресурсов, Дагестанский научный центр Российской академии наук, Махачкала<country>Россия</country></aff><aff xml:lang="en">Institute of Ecology and Sustainable Development, Dagestan State University, Makhachkala; Dagestan Scientific Center, Russian Academy of Sciences, Makhachkala<country>Russian Federation</country></aff></aff-alternatives><aff-alternatives id="aff-2"><aff xml:lang="ru">Институт экологии и устойчивого развития, Дагестанский государственный университет, Махачкала<country>Россия</country></aff><aff xml:lang="en">Institute of Ecology and Sustainable Development, Dagestan State University, Makhachkala<country>Russian Federation</country></aff></aff-alternatives><pub-date pub-type="collection"><year>2016</year></pub-date><pub-date pub-type="epub"><day>07</day><month>10</month><year>2016</year></pub-date><volume>11</volume><issue>3</issue><fpage>35</fpage><lpage>89</lpage><permissions><copyright-statement>Copyright &amp;#x00A9; Абдурахманов Г.М., Набоженко М.В., Абдурахманов А.Г., Иванушенко Ю.Ю., Даудова М.Г., 2016</copyright-statement><copyright-year>2016</copyright-year><copyright-holder xml:lang="ru">Абдурахманов Г.М., Набоженко М.В., Абдурахманов А.Г., Иванушенко Ю.Ю., Даудова М.Г.</copyright-holder><copyright-holder xml:lang="en">Abdurakhmanov G.M., Nabozhenko M.V., Abdurakhmanov A.G., Ivanushenko Y.Y., Daudova M.G.</copyright-holder><license license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>This work is licensed under a Creative Commons Attribution 4.0 License.</license-p></license></permissions><self-uri xlink:href="https://ecodag.elpub.ru/ugro/article/view/856">https://ecodag.elpub.ru/ugro/article/view/856</self-uri><abstract><p>Цель. Целью работы является анализ географических связей Tenebrionidae, результат которого должен лечь в основу фауногенеза Тетийской области Палеарктики. Методы. В основу работы был положен материал, собранный авторами в различных республиках и областях Кавказа, юга европейской части России, Казахстана и Северной Африки за 50 лет (с 1961 по 2016 год), а также обширные литературные данные. Для анализа сходства фаун Тетийской области использовался кластерный анализ на основе коэффициента сходства Жаккара, учитывающий положительные совпадения. Инвазивные виды не включались в анализ. Результаты. Дан обширный обзор истории изучения Tenebrionidae Западной Палеарктики с середины XIX века до наших дней. Намечены основные направления современных исследований. В Тетийской области распространены 403 рода чернотелок, включая лесные группы тропического происхождения, транспалеарктические и космополитные виды, что составляет 63% от всех известных палеарктических родов (640 родов) тенебрионид. Три четверти из них (304 рода, или 75%), объединяющих 5674 вида, составляют преимущественно ксерофильные роды, не встречающиеся за пределами области или представленные лишь очень небольшим количеством видов из пограничных с ней регионов. Доказательства общего исторического развития тенебрионидофауны проявляются в количестве 24 общих (тетийских)   родов, широко распространенных в Тетийской области или немного выходящих за ее границы, но имеющих очень высокое видовое и подродовое разнообразие (вероятно, и центр происхождения) в пределах именно этого региона. Cущественная доля (61%) общететийских родов, высокое разнообразие крупных палеарктических родов Tenebrionidae в западной и восточной частях Тетийской области, а также наличие комплекса супралиторальных реликтов древних морей (Западный Тетис и Восточный Паратетис) из рода Ectromopsis свидетельствуют о единой области формирования фауны чернотелок,    исторически    связанной    с    эпиконтинентальным    палеогеновым    океаном    Тетис.    Выделяется 2 суперрегиона  (с западно-тетийским и восточно-тетийским комплексами таксонов), каждый из которых разделен на несколько субрегонов, отличающихся высоким уровнем таксономического разнообразия. Это деление основано на анализе количества видов и родов, а также уровне эндемизма Tenebrionidae. Наибольшее родовое, видовое разнообразие, а также количество эндемиков отмечается в Северной Африке и Средней Азии. Кластерный анализ общих для области родов, тетийских родов, общих для области видов и рода Blaps на всех дендрограммах показывает сходство фаун Анатолии и Балкан, Кавказ сходет по видовому и родовому составу с Ираном. На всех дендрограммах с большей или меньшей четкостью выделяются два выделенных ранее супперрегиона. При анализе тетийских родов Аравийский полуостров и Северная Африка имеют древние связи с двумя суперрегионами, а при анализе всех видов Тетийской области Арабский полуостров и Ближний Восток имеют древние связи с западно-тетийской и восточно-тетийской фаунами. Заключение. Тенебрионидофауна ряда регионов Тетийской области формировалась по островному типу. Анализ распределения и таксономического разнообразия чернотелок свидетельствует о длительном развитии тенебрионидофаун прибрежных и островных экосистем океана Тетис, а впоследствии независимом расцвете фауны тенебрионид в различных районах тетийского пустынно-степного пояса по мере отступления вод океана, аридизации и орогенеза. При этом в островных горных лесах Анатолии и Атласа с позднетретичной флорой сохранились реликтовые элементы палеогенового происхождения.</p></abstract><trans-abstract xml:lang="en"/><kwd-group xml:lang="ru"><kwd>Западная Палеарктика</kwd><kwd>Tenebrionidae</kwd><kwd>история изучения</kwd><kwd>Тетис</kwd><kwd>Восточный Паратетис</kwd><kwd>сходство и различие фаун</kwd></kwd-group><kwd-group xml:lang="en"><kwd>Western Palaearctic</kwd><kwd>Tenebrionidae</kwd><kwd>history of study</kwd><kwd>Tethys</kwd><kwd>Easten Paratethys</kwd><kwd>similarity and differences of faunas</kwd></kwd-group></article-meta></front><back><ref-list><title>References</title><ref id="cit1"><label>1</label><citation-alternatives><mixed-citation xml:lang="ru">Kergoat G.L., Bouchard P., Clamens A.L., Abbate J.L., Jourdan H., Jabbour-Zahab R., Genson G., Soldati L., Condamine F.L. Cretaceous environmental changes led to high extinction rates in a hyperdiverse beetle family // BMC Evolutionary Biology. 2014. 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